Distribution: Madagascar, adjacent islands. [Lang 1991: map]
Review: Raselimanana 2003: all species. • Descriptions, maps, photos to all species: Glaw & Vences 2007 (Field Guide...)
Zonosaurus aeneus (Grandidier, 1872)
1872 Gerrhosaurus aeneus Grandidier
1942 Zonosaurus aeneus – Angel
Grandidier, A. (1872): Description de quelques reptiles nouveaux découverts a Madagascar en 1870. — Annales des Sciences Naturelles (Paris), (5) 15 (Art. 20): 6–11. [as Gerrhosaurus aeneus; type locality: “Madagascar”; type in MHNP]
Andreone, Glaw, Nussbaum, Raxworthy, Vences & Randrianirina (2003): distr (Nosy Be) • Angel 1942: morph, sys, distr • Brygoo 1985[a]: sys, type in MHNP • Brygoo 1985[b]: all data • Glaw & Vences 1994: distr, morph • Lang 1990[a]: phylogenetic relationships • Meier 1989: morph, distr, behav, habitat, ter • Mocquard 1895[a]: morph • Raselimanana, Raxworthy & Nussbaum 2000: morph, distr, sys • Rogner 1994: morph • Vences, Müller-Jung, Glaw et al. 1996: morph, sys, habitat, distr • Vences, Ziegler, Visser et al. 1999: zoogeography, genital morphology • Wermuth 1968: sys, distr •
Zonosaurus anelanelany Raselimanana, Raxworthy & Nussbaum, 2000
Raselimanana, A. P., C. J. Raxworthy & R. A. Nussbaum (2000): A Revision of the Dwarf Zonosaurus Boulenger (Reptilia: Squamata: Cordylidae) from Madagascar, including Descriptions of Three New Species. — Scientific Papers, Natural History Museum, The University of Kansas, 18: 1–16. [type locality: “Sainte-Luce strand forest, 24°49’S, 47°09’E, 20m elevation, Tolagnaro Fivondronana, Toliara Province, Madagascar”; holotype in UMMZ (= Museum of the Zoology, University of Michigan)]
Zonosaurus bemaraha Raselimanana, Raxworthy & Nussbaum, 2000
Raselimanana, A. P., C. J. Raxworthy & R. A. Nussbaum (2000): A Revision of the Dwarf Zonosaurus Boulenger (Reptilia: Squamata: Cordylidae) from Madagascar, including Descriptions of Three New Species. — Scientific Papers, Natural History Museum, The University of Kansas, 18: 1–16. [type locality: “Antranopasasy, Bemaraha National Park, 18°41.5’S, 44°43.0’E, 100 m elevation, Antsalova Fivondronana, Toliara Province, Madagascar”; holotype in UMMZ (= Museum of the Zoology, University of Michigan)]
Zonosaurus boettgeri Steindachner, 1891
1900 Zonosaurus longicaudatus Mocquard
1942 Zonosaurus boettgeri – Angel
Steindachner, F. (1891): Über einige neue und seltene Reptilien- und Amphibien-Arten. — Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien (Wien), Math. Natur.-wiss. Kl., Abt. 1, 1891: 199–314. [type locality: “Nossi Bé”; type in Naturhist. Museum, Wien]
Andreone, Glaw, Nussbaum, Raxworthy, Vences & Randrianirina (2003): distr (Nosy Be) • Angel 1942: morph, sys, distr • Brygoo 1985[a]: sys about type of Zonosaurus longicaudatus in MHNP • Brygoo 1985[b]: all data • Glaw & Vences 1994: distr, morph • Lang 1990[a]: phylogenetic relationships • Mocquard 1900[a]: description of Zonosaurus longicaudatus (type locality: “Provient d’Andoarano”, type in MHNP) • Mocquard 1900[b]: description (same as above) of Zonozaurus (sic) longicaudatus • Mocquard 1902: sys • Raselimanana, Nussbaum & Raxworthy 2006: observations, re-description • Tiedemann, Häupl & Grillitsch 1994: holotypus in Naturhist. Museum, Wien • Wermuth 1968: sys, distr •
Zonosaurus brygooi Lang & Böhme, 1989
Lang, M. & Böhme, W. (1989): A new species of the Zonosaurus rufipes-complex (Reptilia: Squamata: Gerrhosauridae), from Northern Madagascar. — Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Biologie, 59: 163–168. [see below] [type locality: “Loucoubé, Nosy-Bé, Madagascar”; type in ZFMK]
Andreone, Glaw, Nussbaum, Raxworthy, Vences &
Randrianirina (2003): distr (Nosy Be) • Bauer, Lang &
Günther 1994: sys,
paratypes in ZMB, SMF, IRSNB • Franzen & Glaw
2007: types
• Glaw & Vences
1994: distr,
morph • Nussbaum, Raxworthy,
Raselimanana et al. 1999: distr • Odierna et al.
(2002): Karyotypes • Raselimanana, Raxworthy
& Nussbaum 2000: morph, sys •
Vences,
Müller-Jung, Glaw et al. 1996: morph, sys, habitat, distr •
Vences,
Ziegler, Visser et al. 1999: zoogeography, genital morphology
•
Zonosaurus
haraldmeieri Brygoo
& Böhme, 1985
1985 Zonosaurus
madagascariensis haraldmeieri Brygoo & Böhme
1989 Zonosaurus haraldmeieri – Meier
Brygoo,
É. & Böhme, W. (1985): Un Zonosaurus nouveau de la région d’Antseranana (=
Diégo-Suarez, Madagascar) (Reptilia: Cordylidae). — Revue
Française d’Aquariologie et Herpétologie, 12 (1) (1985):
31–32. [as Zonosaurus madagascariensis
haraldmeieri;
type locality: “Madagascar, Joffreville”; type in ZFMK]
Brygoo 1985[b]:
all data •
Glaw &
Vences 1994: distr,
morph • Henkel & Schmidt
1995: distr, habitat,
size, morph, behav, ter • Lang 1990[a]:
phylogenetic
relationships • Meier 1989:
morph, distr, behav, habitat,
ter • Nietzke 1998:
ter, morph, distr, behav,
diet, reproduction • Rogner 1991[a]:
ter, breeding •
Rogner
1994: morph, behav,
ter •
Zonosaurus
karsteni (Grandidier,
1869)
1869
Gerrhosaurus karsteni Grandidier
1942 Zonosaurus karsteni – Angel
Grandidier, A.
(1869): Descriptions
de quelques animaux nouveaux découverts, pendant l’année
1869, sur la côte ouest de Madagascar. — Revue et Magazine
de Zoologie (Paris), (Sér. 2) 21: 337–342. [as
Gerrhosaurus
karsteni; type
locality: “Fiérin”; syntype in MHNP]
Angel 1942:
morph, sys, distr
• Boettger 1913:
morph •
Böhme
1988: genital
morphology • Breustedt 1990:
ter, breeding •
Brygoo
1985[a]: sys, syntype
in MHNP • Brygoo 1985[b]:
all data •
Glaw &
Vences 1994: distr,
morph • Henkel & Schmidt
1995: distr, habitat,
size, morph, behav, ter • Lang 1990[a]:
phylogenetic
relationships • Meier 1989:
morph, distr, behav, habitat,
ter • Mocquard
1895[a]: morph
• Mocquard
1900[a]: morph
• Odierna et al.
(2002): Karyotypes • Rogner 1994:
morph, behav, ter
• Wermuth 1968:
sys, distr
•
Zonosaurus
laticaudatus (Grandidier,
1869)
1869
Gerrhosaurus laticaudatus Grandidier
1942 Zonosaurus laticaudatus – Angel
Grandidier,
A. (1869): Descriptions de quelques animaux nouveaux
découverts, pendant l’année 1869, sur la côte ouest de
Madagascar. — Revue et Magazine de Zoologie (Paris), (Sér.
2) 21: 337–342. [as Gerrhosaurus
laticaudatus;
type locality: “Fiérin”; holotype in Naturhist. Museum,
Wien]
Andersson 1910:
morph •
Andreone,
Glaw, Nussbaum, Raxworthy, Vences & Randrianirina
(2003): distr (Nosy
Be) • Angel 1942:
morph, sys, distr
• Angel 1949:
morph •
Boettger
1913: morph
• Bowler 1977:
age: >8 years
• Brygoo 1985[a]:
sys, syntype in MHNP
• Brygoo 1985[b]:
all data •
D’Cruze
2006: semiaquatic
defensive behaviour • Glaw & Vences
1994: distr,
morph • Glos 2004:
feeding on frogs
• Harvey & Gutberlet
1995: microstructure,
evolution and ontogeny of scale surfaces •
Henkel
& Schmidt 1995: distr, habitat, size, morph, behav, ter
• Hoesch 1985:
morph •
Kaudern
1922: morph
• Lang 1990[a]:
phylogenetic
relationships • Meier 1989:
morph, distr, behav, habitat,
ter • Mertens 1970:
age: >8 years
• Mocquard
1895[a]: morph
• Nussbaum, Raxworthy,
Raselimanana et al. 1999: distr • Odierna et al.
(2002): Karyotypes • Putti & Della Rossa
1996: cytology
• Randriamahazo
1998: activity
temperatures • Rogner 1994:
morph, behav, ter
• Snider & Bowler
1992: age: >8 years
• Tiedemann, Häupl &
Grillitsch 1994: holotype in Naturhist.Museum, Wien
• Urbani & Bels
1995: description of
feeding phases • Wermuth 1968:
sys, distr
•
Zonosaurus
madagascariensis madagascariensis (Gray,
1831)
1831
Cicigna madagascariensis Gray
1839 Gerrhosaurus bifasciatus Duméril & Bibron
1985 Zonosaurus madagascariensis madagascariensis
– Brygoo
Gray, J.
(1831): A Synopsis of
the Species of the Class Reptilia. — In: Griffith, Edward
& Pidgeon, Edward (Hrsg.): The Animal Kingdom arranged
in conformity with its Organisation by the Baron Cuvier
[...]. [9] – London: Whittaker, Treacher & Co., 1–110.
[as Cicigna
madagascariensis;
type locality: “Madagascar”; type in
NHM]
Andreone, Glaw,
Nussbaum, Raxworthy, Vences & Randrianirina
(2003): distr (Nosy
Be) • Andreone &
Randriamahazo 1997: distr, morph (Z. cf. madagascariensis)
• Angel 1942:
morph, sys, distr
• Angel 1949:
morph •
Bissinger
& Simon 1979: tongue extrusions •
Böhme
1988: genital
morphology • Boulenger
1887[a]: morph
• Broadley
1971[b]: morph,
distr • Brygoo 1985[a]:
sys, syntype of
Gerrhosaurus
bifasciatus in
MHNP • Brygoo 1985[b]:
all data •
Dumeril
& Bibron 1839: description of Gerrhosaurus
bifasciatus,
type locality: “Madagascar” • Gehring 2006:
predation of
Paroedura
• Glaw & Vences
1994: distr,
morph • Gray 1845:
morph •
Henkel
& Schmidt 1995: distr, habitat, size, morph, behav, ter
• Kaudern 1922:
morph •
Lang
1990[a]: phylogenetic
relationships • Meier 1989:
morph, distr, behav, habitat,
ter • Methuen & Hewitt
1913: morph
• Miehle & Bauer
1993: visceral
vasculature • Morescalchi et al.
1997: regulatory
peptides, serotonin, gut, pancreas,
immunohistochemistry • Odierna et al.
(2002): Karyotypes • Rogner 1994:
morph •
Schmidt
1913: skin, dermal
bones • Wermuth 1968:
sys, distr •
Wever
1978: structure and
function of ear •
Zonosaurus
madagascariensis insulanus Brygoo,
1985
Brygoo, É.
(1985[b]): Les
Gerrhosaurinae de Madagascar, Sauria (Cordylidae). —
Mémoires du Muséum National d’Histoire Naturelle, Paris,
sér. A, Tome 134: 1–65. [type locality: “Glorioso Islands
NW Madagascar”; holotype in NHM]
Zonosaurus
maramaintso Raselimanana,
Nussbaum & Raxworthy, 2006
Raselimanana, Achille
P., R. A. Nussbaum & C. J. Raxworthy (2006):
Observations and
re-description of Zonosaurus boettgeri
Steindachner 1891 and
description of a second new species of long-tailed
Zonosaurus
from western Madagascar. —
Occasional Papers of the Museum of Zoology, University of
Michigan, August 2006, No. 739: 1-16. [type locality:
“Antsalova region, Antsalova Fivondronana, Mahajanga
Province, Madagascar”, holotype in UMMZ, No. 221904 an
adult male]
[Love] 2008:
nice photo at the cover of
Herpetological Review, 39(2) [see above; thanks to Robert
Hansen] •
Zonosaurus
maximus Boulenger,
1896
Boulenger, G.
(1896[a]): Descriptions of new Lizards from
Madagascar. — The Annals and Magazine of Natural History
(London), (6), Vol. 17: 444–449. [type locality: “Imerina,
Mad.”; type in NHM]
Angel 1942:
morph, sys, distr
• Blanc 1967:
behav, morph •
Böhme
1988: genital
morphology • Brygoo 1985[b]:
all data •
Glaw &
Vences 1994: distr,
morph • Harvey & Gutberlet
1995: microstructure,
evolution and ontogeny of scale surfaces •
Henkel
& Schmidt 1995: distr, habitat, size, morph, behav,
ter • Lang 1990[a]:
phylogenetic
relationships • Meier 1989:
morph, distr, behav, habitat,
ter • Rogner 1994:
morph, behav, ter
• Wermuth 1968:
sys, distr •
Zonosaurus
ornatus (Gray,
1831)
1831
Cicigna madagascariensis var. ornata Gray
1834 Gerrhosaurus lineatus Cocteau
1887 Zonosaurus ornatus – Boulenger
Gray, J.
(1831): A Synopsis of
the Species of the Class Reptilia. — In: Griffith, Edward
& Pidgeon, Edward (Hrsg.): The Animal Kingdom arranged
in conformity with its Organisation by the Baron Cuvier
[...]. [9] – London: Whittaker, Treacher & Co., 1–110.
[as Cicigna
madagascariensis var. ornata; type locality: “Madagascar”; type in
NHM]
Angel 1942:
morph, sys, distr
• Boulenger
1887[a]: morph
• Brygoo 1985[a]:
sys, type of
Gerrhosaurus
lineatus in MHNP
• Brygoo 1985[b]:
all data •
Cocteau
1834: description
of Gerrhosaurus
lineatus,
type locality: “Madagascar” • Glaw & Vences
1994: distr,
morph • Gray 1845:
morph •
Harvey
& Gutberlet 1995: microstructure, evolution and ontogeny of
scale surfaces • Henkel & Schmidt
1995: distr, habitat,
size, morph, behav, ter • Lang 1990[a]:
phylogenetic
relationships • Meier 1989:
morph, distr, behav, habitat,
ter • Methuen & Hewitt
1913: morph
• Nguyen-N-Anh
1974: l’ultrastructure
du cone papillaire • Odierna et al.
(2002): Karyotypes • Rogner 1994:
morph •
Wermuth
1968: sys, distr
• Wever 1978:
structure and function of ear
•
Zonosaurus
quadrilineatus (Grandidier,
1867)
1867
Gerrhosaurus 4-lineatus Grandidier
1942 Zonosaurus quadrilineatus – Angel
Grandidier,
A. (1867): Liste des
reptiles nouveaux découverts, en 1866, sur la côte
sud-ouest de Madagascar. — Revue et Magazine de Zoologie
(Paris), (Sér. 2) 19: 232–234. [as Gerrhosaurus
4-lineatus; type
locality: “Tullear”; type in MHNP]
Angel 1942:
morph, sys, distr
• Boettger 1913:
morph •
Böhme
1988: genital
morphology • Brygoo 1985[a]:
sys, type in MHNP
• Brygoo 1985[b]:
all data •
Glaw &
Vences 1994: distr,
morph • Henkel & Schmidt
1995: distr, habitat,
size, morph, behav, ter • Lang 1990[a]:
phylogenetic
relationships • Meier 1988:
morph, distr, behav, habitat,
ter • Mocquard
1895[a]: morph
• Rogner 1994:
morph, behav, ter
• Tiedemann, Häupl &
Grillitsch 1994: syntype in Naturhist. Museum, Wien
• Wermuth 1968:
sys, distr •
Zonosaurus
rufipes (Boettger,
1881)
1881
Gerrhosaurus (Cicigna) rufipes Boettger
1887 Zonosaurus rufipes – Boulenger
Boettger,
O. (1881[a]): Diagnoses reptilium et batrachiorum
novorum ab ill. Antonio Stumpff in insula Nossi-Bé
Madagascariensi lectorum. — Zoologischer Anzeiger
(Leipzig), 4 (87): 358–362. [as Gerrhosaurus (Cicigna)
rufipes; type
locality: “Nossi Bé”; type in SMF]
Andreone, Glaw,
Nussbaum, Raxworthy, Vences & Randrianirina
(2003): distr (Nosy
Be) • Angel 1942:
morph, sys, distr
• Bauer, Lang &
Günther 1994: sys,
paralectotypes in ZMB • Boulenger
1887[a]: morph
• Brygoo 1985[b]:
all data •
Glaw &
Vences 1994: distr,
morph • Henkel & Schmidt
1995: distr, habitat,
size, morph, behav, ter • Lang 1990[a]:
phylogenetic
relationships • Lang & Böhme
1989: sys
• Meier 1989:
morph, distr, behav, habitat,
ter • Mertens 1922:
type in SMF •
Mertens
1967: type in SMF
• Odierna et al.
(2002): Karyotypes • Rogner 1994:
morph •
Raselimanana, Raxworthy
& Nussbaum 2000: morph, sys • Tiedemann, Häupl &
Grillitsch 1994: paralectotypes in Naturhist. Museum,
Wien • Vences, Müller-Jung,
Glaw et al. 1996: morph, sys, habitat, distr •
Wermuth
1968: sys, distr
•
Zonosaurus
subunicolor (Boettger,
1881)
1881
Gerrhosaurus (Cicigna) rufipes var. subunicolor
Boettger
1994 Zonosaurus subunicolor – Glaw & Vences
1996 Zonosaurus subunicolor – Vences, Müller-Jung,
Glaw & Böhme
Boettger,
O. (1881[a]): Diagnoses reptilium et batrachiorum
novorum ab ill. Antonio Stumpff in insula Nossi-Bé
Madagascariensi lectorum. — Zoologischer Anzeiger
(Leipzig), 4 (87): 358–362. [as Gerrhosaurus (Cicigna)
rufipes var. subunicolor; type locality: “Nossi Bé”; type in
SMF]
Andreone, Glaw,
Nussbaum, Raxworthy, Vences & Randrianirina
(2003): distr (Nosy
Be) • Angel 1942:
morph, sys, distr
• Glaw & Vences
1994: distr,
morph • Lang & Böhme
1989: sys
• Mertens 1922:
type in SMF •
Mertens
1967: type in SMF
• Raselimanana, Raxworthy
& Nussbaum 2000: morph, sys • Vences, Müller-Jung,
Glaw et al. 1996: morph, sys, habitat, distr •
Zonosaurus
trilineatus Angel,
1939
Angel, F.
(1939): Un
Gerrhosauridé nouveau de Madagascar. — Bulletin de la
Société Zoologique de France (Paris), 64: 350–351. [type
locality: “Ambovombe (Sud de Madagascar)” ; type in
MHNP]
Angel 1942:
morph, sys, distr
• Brygoo 1985[a]:
sys, abou type in MHNP
• Brygoo 1985[b]:
all data •
Glaw &
Vences 1994: distr,
morph • Henkel & Schmidt
1995: distr, habitat,
size, morph, behav, ter • Lang 1990[a]:
phylogenetic
relationships • Odierna et al.
(2002): Karyotypes • Rogner 1994:
morph •
Wermuth
1968: sys, distr
•
Zonosaurus
tsingy Raselimanana,
Raxworthy & Nussbaum, 2000
Raselimanana, A. P., C.
J. Raxworthy & R. A. Nussbaum (2000):
A Revision of the
Dwarf Zonosaurus Boulenger (Reptilia: Squamata:
Cordylidae) from Madagascar, including Descriptions of
Three New Species. — Scientific Papers, Natural
History Museum, The University of Kansas, 18: 1–16. [type
locality: “Lac Vert, Ankarana National Park, 12°52’S,
49°06’E, 100 m elevation, Ambilobe Fivondronana,
Antsiranana Province, Madagascar”; holotype in UMMZ (=
Museum of the Zoology, University of
Michigan)]
© Klaus Adolphs – created: 26 February 2001 – last changed or updated: 27 December 2009

